Descriptions of Subgenera:
- Subgenus Heliacus (Heliacus) ORBIGNY in SAGRA, 1842
Teleoconch: small to medium-sized (usually 6-13 mm, rarely over 20 mm), depressed to tall roundly cone-shaped, with narrow to very wide umbilicus (ca. 8-40% of shell diameter); whorls weakly to distinctly bulging; sculpture: spiral ribs with relatively large nodules on entire surface; number of ribs constant, only between lower peripheral and infraperipheral ribs occasionally with 1 ± strong additional rib; apical side: subsutural rib, single midrib (or two narrow ones) and upper peripheral rib usually equally developed, rarely upper peripheral rib somewhat stronger; periphery formed by lower peripheral rib and only slightly weaker infraperipheral rib (occasionally with additional rib between them); upper point of whorl attachment at or below lower peripheral rib; base usually with 5 spiral ribs, increasing in width towards umbilicus; umbilical wall usually with 1-2 strong, narrow spiral ribs; coloration of (in most cases distinctly demarcated) brown to black flecks and flames on ± white background, with peripheral ribs as a rule with very regular pattern; subsutural rib, infraperipheral rib and rib surrounding umbilicus (UC) often lighter colored. Protoconch: very small to medium-sized (0.55-1.05), distinctly heterostrophic, without anal keel- Radula: five-toothed-taenioglossate; rachidian with strong central cusp flanked by numerous (12-20) smaller cusps on either side; inner and outer marginals with 5-13 cusps each. Operculum: horny, round, cone-shaped, with peg-like projection on body side.
- Subgenus Heliacus (Pyrgoheliacus) BIELER, 1987
Teleoconch: very small to small (usually 2.6 to 8 mm), tall cone-shaped (in large specimens, height ± equals diameter), with narrow to moderately wide umbilicus (ca. 10-20% of shell diameter); whorls distinctly bulging, periphery rounded; sculpture: spiral ribs with ± rounded nodules on entire surface; apical side: subsutural rib, and 1-2 midribs of about equal strength, upper peripheral rib more prominent and sometimes (turritus) already part of the periphery; periphery otherwise formed by ± equally strong lower peripheral rib (most prominent part of periphery), strong infraperipheral rib, and an almost equally strong additional rib between them (here upper point of whorl attachment); suture distinct; base with 6 spiral ribs, increasing in width towards umbilicus, innermost somewhat sunken into umbilicus; umbilical wall bulging, with or without weak spiral threads; coloration off-white to yellowish-tan; peripheral ribs with pattern of brown flecks (pattern corresponding between various ribs); inner lip off-white to dark brown. Protoconch; small to medium-sized (0.82-0.98), distinctly heterostrophic, with or without anal keel. Radula: five-toothed taenioglossate; rachidian with strong central cusp flanked by smaller cusps on either side; inner and outer marginals with several cusps each (Atlantic H verdensis). Operculum: horny, round, cone-shaped, with peg-like projection on body side.
- Subgenus Heliacus (Torinista) IREDALE, 1936
Teleoconch; very small to medium-sized (usually 3-12 mm), in most cases disk-shaped with weakly bulging whorls, occasionally depressed cone-shaped; umbilicus very narrow to extremely wide (ca. 9-55% of shell diameter); sculpture: spiral ribs with ± fine nodules, often flattened on the midribs; distinct axial sculpture in forms with fused spiral ribs; apical side: subsutural rib usually somewhat wider than following (often fused) midrib; periphery formed by somewhat stronger lower peripheral rib and infraperipheral rib, between them often 1-2 ± strong additional ribs; upper point of whorl attachment at lower peripheral rib, infraperipheral rib, or between them; base usually with 6 spiral ribs, increasing in width towards umbilicus; umbilical wall smooth except for axial growth lines, or with 1-2 ± demarcated spiral ribs; coloration of usually irregular flames in various shades of brown on apical and basal sides, only peripheral ribs often with regular pattern of light and dark brown flecks; innermost rib on base (UC) in most cases light in color. Protoconch: small to large (0.65-1.40), distinctly heterostrophic, in most cases with several folds in false umbilicus, rarely with anal keel. Radula: five-toothed taenioglossate; rachidian with strong central cusp flanked by numerous (6-20) smaller cusps on either side; inner and outer marginals with 6-10 cusps each. Operculum; horny, round, cone-shaped (often taller than diameter), with peg-like projection on body side.
- Subgenus Heliacus (Grandeliacus) IREDALE, 1957
Teleoconch: shell small to large (usually 9-33 mm); small shells with angular periphery and basal area more convex than apical side; larger shells with rounded periphery and bulging whorls; umbilicus moderately wide to wide (ca. 17-36% of shell diameter); sculpture: on early whorls, subsutural rib and 2 midribs ± weakly developed and upper peripheral rib prominent (concave area between upper and lower peripheral rib); on later whorls, subsutural rib stronger, or subsutural rib, midribs and upper peripheral rib nearly equally strong (often additional rib between upper and lower peripheral rib); periphery: early whorls two-ribbed, with somewhat more prominent lower peripheral rib and infraperipheral rib; later, 1-3 additional spiral ribs between them, one of which often equally prominent; upper point of whorl attachment at lower peripheral or additional rib; base with 6 (later 7) spiral ribs, increasing in width towards umbilicus; large specimens with additional finer ribs interspersed; umbilical wall usually with 1-2 fine spiral ribs; coloration yellowish tan, early whorls often darker; peripheral ribs sometimes with fleck pattern. Protoconch: small to medium-sized (0.78-1.20), distinctly heterostrophic, without anal keel. Radula: five-toothed-taenioglossate; rachidian with central cusp flanked by numerous (9-22) cusps on either side, with central cusp larger than or equal to flanking cusps; inner and outer marginals with 6-10 cusps each. Operculum; horny, round, flat in young individuals, later moderately cone-shaped, with peg-like projection on body side.
- Subgenus Heliacus (Teretropoma) ROCHEBRUNE, 1881
Teleoconch: small to medium-sized (usually 5.5-13 mm); depressed to turreted cone-shaped, always with distinctly rounded whorls; young or depressed shells with two-ribbed periphery; umbilicus moderately wide to extremely wide (23-65% of shell diameter); sculpture: axial growth sculpture on entire surface; apical side: subsutural rib, single midrib, and upper peripheral rib ± equally strong developed, at least in fully grown specimens interspersed with finer spiral threads; periphery formed by almost equally strong lower peripheral and infraperipheral ribs, always flanked by additional weaker ribs; upper point of whorl attachment at infraperipheral rib under formation of a usually deep suture; base with ca. 6 stronger spiral ribs (with usually one weaker additional rib between them); basal area angular or rounded at umbilicus; with spiral sculpture on the umbilical wall; white, solid brown, or with ± irregular flames of tan or reddish to dark brown, with periphery and umbilicus often lighter. Protoconch: small to large (0.72-1.24), distinctly heterostrophic, without anal keel. Radula: five-toothed-taenioglossate; rachidian with strong central cusp flanked by numerous (9-14) smaller cusps on either side; inner and outer marginals with about 7 cusps each. Operculum: horny, round, moderately cone-shaped or flat to concave, with peg-like projection on body side.
- Subgenus Heliacus (Gyriscus) TIBERI, 1867
Teleoconch: small (usually 6-10 mm); tall, roundly cone-shaped with distincdy rounded, bulging whorls; umbilicus very narrow to narrow (ca. 5-10% of shelldiameter); entire surface with fine granular sculpture formed by crossing fine axial growth threads and narrow spiral ribs; later whorls with many additional spiral ribs which are almost as strong as the main ribs; apical side: subsutural rib, two midribs, upper peripheral rib, and several additional ribs interspersed; broad, rounded periph¬eral region demarcated by lower peripheral and infraperipheral rib, with 1-4 ± strong and sometimes numerous weaker, additional ribs between them; upper point of whorl attachment between upper and infraperipheral ribs, at additional ribs of periphery; no distinct suture; base with 6-8 stronger and numerous finer spiral ribs; umbilical wall usually with 1-2 ± irregular spiral ridges, in large specimens often smooth except for growth lines; usually white, occasionally with weak tan flames and brown parietal wall. Protoconch: medium-sized (0.90-1.15), distincdy heterostrophic, protruding bubble-like, without anal keel. Radula: five-toothed-taenioglossate; rachidian with strong central cusp flanked by numerous (10-12) smaller cusps on either side; inner and outer marginals with 4-8 cusps each [Mediterranean H jeffreysianus]. Operculum; horny, round, weakly cone-shaped, with peg-like projection on body side.