Tegula Lesson, 1832 Tegula Lesson, 1832 [1832-1835]: pl. 51. Type species (M): Tegula elegans Lesson, 1832 [= Trochus pellisserpentis Wood, 1828]. Panama. Agathistoma Olsson & Harbison, 1971: 350. Type species (OD) Trochus viridulus Gmelin, 1791. Caribbean. Promartynia Dall, 1909: 94. Type species (OD): Trochus pulligo Gmelin, 1791. Northeast Pacific. Diagnosis. Shell small to medium in size, sturdy with dome-like spire or coniform; periphery sharply angulate or rounded; sculpture smooth, spiral, or rugose. Umbilicus open in juvenile shells, opened in mature specimens or covered with callus, leaving broad umbilical depression. Columellar wall arched, columella with denticles, largest one at termination of spiral cord bordering or passing into umbilical cavity; aperture oblique, sometimes with lirations inside, interior nacreous, white or brightly colored. Operculum uncalcified. Color always dark, usually gray, black, or brown. Remarks. The genus name Chlorostoma was used for the West American Tegula-species by different authors in the past (see below). Williams et al. (2008) showed that Tegula and Chlorostoma are well separated and Chlorostoma is only available for 3 species form the West Pacific.