Habitat: it lives in the infra-circalittoral zones, into substratum cavities and at the margins of Posidonia oceanica meadows. Distribution: it can be found in a large part of the Mediterranean but never common. More frequent in the Southern area. Imaged shells: even in its yuvenile stage, this species cannot be conftised with any other Mediterranean species. Shell solid, slim in form, bright, wholly smooth. Upper spiral elevated, apex smooth and obtuse. Last whorl is almost the totality of its height, suture superficial and not much distinguishable. Aperture narrow and long. External lip without teeth and thickened, with a slight flexuosity in the meridian part, in the upper area it can be noted one slight shoulder. On columella there are four quite clear folds.
Fresh specimens are usually either yellow or light brown in colour, sometimes either with little bands or maculae brown in colour. This species has no operculum. It is easily recognizable from Gibberula, due to its bigger sizes and to its upper spiral which is much more elevated. Juvenile specimens are more roundish in outline. Gofas (1989) considers Marginella calameli Jousseaume, 1875, species collected along Algerian coasts having one shorter and rolling spiral and two darker spiral bands, as a simple variant of V. mitrella. The adult specimens average measures are around 8-10 mm in height.