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Taxon profile

species

rough periwinkle
Littorina saxatilis (Olivi, 1792)

kingdom Animalia - animals »  phylum Mollusca - mollusks »  class Gastropoda - gastropods »  order Littorinimorpha »  family Littorinidae - Periwinkles »  genus Littorina

Scientific synonyms

Littorina saxatilis saxatilis (Olivi, 1792)
Littorina rudis Maton, 1797

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Description

Diagnostic characters
Shell solid, opaque, with pointed spire, swollen whorls and deep sutures; ornament of numerous spiral ridges which are usually symmetrically V-shaped in section; examine those not on shell base, and see below), sometimes reduced and often eroded. Aperture round, outer lip arising from last whorl almost at right angles to shell axis. Columella short, apertural edge turned out below it slightly. Colour variable, but usually with a tesselated pattern, throat darker than outer surface. Tip of penis often blunt but with a point, its glands small and not reaching the curved base. Ovoviviparous. terminal part of the female duct containing embryos.
Other characters
The shell has 4-5 whorls. The ratio breadth of last whorl/breadth of penult is two or more (breadth = greatest measurement at right angles to shell axis; the breadth of the last whorl measured just above the origin of the outer lip). The area of mantle skirt between the genital duct and the columellar muscle may show red colour in live animals (it is sometimes difficult to be sure of this) and in males the prostate gland extends anterior to the anus. Up to 15 mm high, 10 mm broad; last whorl occupies 75-80% of shell height, aperture 50-60%.
L. saxatilis, the highest littorinid apart from L. neritoides, is typically found in crevices of bedrock at and above the Pelvetia zone. The species is spread throughout western Europe, including the western parts of the Baltic, and is also found on north eastern and north western coasts of North America. A dwarf form with large aperture. L. s. scotia, has been described from algal tufts in Rockall, and another with everted lips (L. s. panda) also from Rockall and Eddystone Rocks probably belongs here.
Some authorities, most recently J. E. Smith (1981), regard the form rudis Matcn, 1797 as a species distinct from saxatilis, though most treat it only as an ecological variant. The differences by which rudis may be recognized are: the spiral ridges are asymmetrical in section, their adapical sides the steeper; the out-turning of the lip below the columella is large: the shell usually shows no tessellated pattern; the throat is not dark; the ratio breadth of last whorl/breadth of penult is 2 or less; the animals occur about the level of Pelvetia canaliculata but prefer damp areas where they may be found under stones; if on vertical surfaces they are not usually in crevices.
Graham, A.; 1988. Molluscs: Prosobranch and Pyramidellid Gastropods.
Author: Jan Delsing

Similar species

Littorina arcana Ellis, 1978

Included taxa

Number of records: 6

subspecies Littorina saxatilis albida Dautzenberg, 1882

Littorina saxatilis albida

subspecies Littorina saxatilis aurantia Dautzenberg, 1882

Littorina saxatilis aurantia

subspecies Littorina saxatilis fusca Dautzenberg & Fischer, 1912

Littorina saxatilis fusca

subspecies Littorina saxatilis lutea Dautzenberg & Durouchoux, 1900

Littorina saxatilis lutea

subspecies Littorina saxatilis spitsbergica Kimchowitz, 1896

Littorina saxatilis spitsbergica

subspecies Littorina saxatilis zonaria W.I. Bean, 1844

Littorina saxatilis zonaria


Links and literature

EN Galli C.: WMSDB - Wolrdwide Mollusc Species Data Base July 10, 2013 [http://www.bagniliggia.it/WMSD/WMSDhome....] [as Littorina saxatilis Olivi, 1792]
Data retrieved on: 23 November 2013
CZ Pfleger V. (1999): České názvy živočichů III. Měkkýši (Mollusca), Národní muzeum, (zoologické odd.), Praha, 108 pp. [as Littorina saxatilis (OLIVI, 1792)]
Data retrieved on: 11 November 2013
IT Repetto G., Orlando F. & Arduino G. (2005): Conchiglie del Mediterraneo, Amici del Museo "Federico Eusebio", Alba, Italy [as Littorina saxatilis (Olivi, 1792)]

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