TYPE MATERIAL - Lectotype, here designated, RGM 607 310a (Koperberg 1931, pl. 3, fig. 35), 3 paralectotypes RGM 607
310b, RGM 607 309a, b (as listed below).
TYPE LOCALITY Indonesia, Timor; Pleistocene.
DISTRIBUTION- Western Pacific, known from Taiwan in the north, Indonesia in the west, to Fiji in the southeast; 96- 810, alive in 96-642 m. Syntopic with M. kuroharai Azuma, 1960 in Taiwan (TAIWAN 2004 stn CP269) and the Philippines (AURORA stn CP2749); with M. clandestina in the Philippines (MUSORSTOM 2 stn CP78; PANGLAO 2005 stn CP2336, CP2351, CP2388.

DESCRIPTION - Shell small for genus, up to 22.2 mm (BORDAU 1), thick, solid, white. Shape slender, fusiform with elongate spire, base convex, siphonal canal moderately short. Protoconch 1.25 smooth, glossy whorls, diameter 0.9 mm. Transition to teleoconch indistinct, marked by first axial rib. Teleoconch of six convex whorls, upper spire whorls angulated. Suture distinct. First teleoconch whorl with six fine spiral cords, interspaces each equal in width to, or twice as wide as, one cord. Both subsutural interspaces becoming wider by intercalation of fine secondary spiral cord on third whorl. Secondary cords gradually increasing in strength, becoming as strong as primary cords on fourth whorl, producing eight spiral cords of equal strength. Ninth cord may be partially concealed under lower suture in few specimens. Spiral cords of most specimens wide, slightly flattened, with interspaces each equal in width to one cord, narrow with wider interspaces in few specimens. Rarely interspaces with single, fine, secondary spiral cord. Spiral cords appearing narrower in specimens with periostracum removed. Penultimate whorl with eight to nine spiral cords. Last whorl with ca. 30 spiral cords, 15 on whorl, ca. 15 on siphonal canal. All spire whorls with 10-11 strongly convex axial ribs, inters- paces each twice as wide as one rib. Axial ribs becoming more angulate on second whorl. Third and fourth whorls angulate. Axial ribs narrower, with wider interspaces, in few specimens. Spiral interspaces crossed by fine incremental lamellae, indistinct in axial interspaces, pronounced on axial ribs. Aperture oval. Columella smooth, glossy, slightly impressed in few specimens. Outer lip thin, thickened in few specimens, with five to seven fine internal lirae or thick knobs. Siphonal canal short, rather narrow, open. Aperture and siphonal canal together slightly less than half of total shell length. Periostracum thin, yellowish-brown, forming accentuated incremental lamellae in axial interspaces. Animal, radula and operculum unknown.

REMARKS― Beets (1953: 246) was the first to recognize the placement of Lathyrus jonkeri in Manaria. Lot RGM 607 309 of the original syntype series is heterogenous and contains a specimen RGM 607 309a) that clearly belongs to Lathyrus jonkeri and a broken shell (RGM 607 309b) belonging to Fusinus. The generic placement suggested by Robba et al. (1989: 80) must have been based on this last specimen. Manaria jonkeri is characterized by a moderately slender shape, rather equally spaced spiral cords, axial ribs that form an angulate shoulder on the upper spire whorls but become adapically slightly weaker on the later whorls, in combination with rather convex penultimate and last whorls when fully adult. Some specimens from the northern Philippines (MUSORSTOM 3 stn CP136 and all listed AURORA 2007 stations) have the aperture ornamented with large knobs on the columella and the inner edge of the outer lip, in combination with an inflated last whorl bearing a higher number of axial ribs. Despite this striking morphological difference, we regard this pronounced sculpture in combination with an inflated last whorl as falling within the range of intraspecific variation of M. jonkeri, based on the identical shape and sculpture of the upper spire whorls. A few specimens from Taiwan have a fine secondary spiral cord in each interspace on the base and most have slightly more angled whorls than the specimens from the Bohol Sea (PANGLAO 2005) but intermediate specimens have been observed. Manaria lirata Kuroda & Habe, 1961 differs from M. jonkeri in the broader shape, the less strongly convex spire whorls, the rather angulate whorls with a wide subsutural concavity, the typically weaker spiral sculpture with wider interspaces, and the angulation on the axial ribs near the periphery (compared to quite adapical in M. jonkeri). Manaria clandestina Bouchet & Warén, 1986 differs from M. jonkeri in its higher spire, sharper axial ribs, slightly wider spiral interspaces, more convex shoulder and larger adult size. Manaria sp. 1 ( has a similar shape to that of M. jonkeri but differs by the finer spiral cords with wider interspaces, the slightly narrower axial ribs and the larger adult size. Adult specimens of M. jonkeri have strong apertural lirae inside the outer lip while juvenile specimens of Manaria sp. 1 have a smooth aperture, but otherwise shells of about the same size of both species are difficult to distinguish. Manaria loculosa n. sp. differs from M. jonkeri in the broader shape, the wider interspaces with a secondary spiral cord in a few specimens, and the angulate axials near the upper suture (instead of near the periphery). Manaria corporosis n. sp. possesses similar sculpture when juvenile but differs from M. jonkeri in the more slender shape, the wider spiral cords with narrower interspaces, the slightly fewer axial ribs, and the more adapical shoulder angulation, with a narrower subsutural slope.