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Kohlmann B, Solís Á, Alvarado G (2019):
    Description of Onthophagus humboldti and Uroxys bonplandi, two new scarab beetles (Coleoptera, Scarabaeidae, Scarabaeinae) from Costa Rica, with notes on tropical mountain brachyptery and endemicity

 
 
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Uroxys bonplandi
BIOLIB: Kohlmann, Solís & Alvarado, 2019
Onthophagus humboldti Kohlmann, Solís & Alvarado, 2019
 
 
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