Grant, T., M. Rada, M. A. Anganoy-Criollo, A. Batista, P. H. dos S. Dias, A. M. Jeckel, D. J. Machado, and J. V. Rueda-Almonacid (2017): Phylogenetic systematics of dart-poison frogs and their relatives revisited (Anura: Dendrobatoidea), South American Journal of Herpetology 12 (Special Issue): 1–90 [as Leucostethus Grant, Rada, Anganoy-Criollo, Batista, Dias, Jeckel, Machado & Rueda-Almonacid, 2017]
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