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Taxon profile

genus

Clanculus Montfort, 1810

kingdom Animalia - animals »  phylum Mollusca - mollusks »  class Gastropoda - gastropods »  family Trochidae - top-shells »  tribus Trochini

Scientific synonyms

incl. Clanculopsis T.A. de M. Monterosato, 1888, Clanculus Montfort, 1810, Paraclanculus Finlay, 1926
Euclanculus Cotton & Godfrey, 1934
Euriclanculus Cotton & Godfrey, 1934
Fragella Swainson, 1840
Isoclanculus Cotton & Godfrey, 1934
Macroclanculus Cotton & Godfrey, 1934
Mesoclanculus Iredale, 1924
Microclanculus Cotton & Godfrey, 1934

Description

The shell is of medium size, turbiniform and with whorls and base slightly convex. The surface is granulose. The parietal lip is crenulated and curved, delimiting a false umbilicus. The columella presents a very evident tooth, simple or complex.
The animal has a well-developed epipodium. The cervical lobes can have the margins finely crenulate, being usually similar on both sides, rarely asymmetrical. The cephalic lappets are large. Four pairs of epipodial tentacles.
The species living in the Mediterranean and West Africa have radular characters similar to other Trochidae of the subfamily Trochinae, tribe Margaritini subgenus Valvatella, and tribe Cantharidini. Their radular formula is: n-5-1-5-n. The rachidian tooth has a very wide base which narrows to a cusp, and presents two basic forms: triangular and ovoid; its cusp is M-shaped, sharp or rounded anteriorly and has 3-5 sharp denticles on either side of the base. Lateral teeth are similar to the rachidian in size and shape; the outermost lateral is a little narrower and stronger. Marginal teeth elongate and curved forming a hook-like angle at the beginning of the cusp.
Rubio, F. & Rolán, E., 2002. Revision of the Genus Clanculus (Gastropoda: Trochidae) in the Eastern Atlantic.

Taxonomy

The animal of Clanculus bears four pairs of tentacular filaments on the epipodial line. The dentition (T. pharaonius) is similar to that of Trochus, but the body of the tooth is more expand¬ed, the centrals and laterals bearing large lateral supporting-wings.
I was at first inclined to treat Clanculus as a genus distinct from Trochus; but on account of the difficulty of so defining the group that its species may always be distinguished from Trochus, and the lack of any tangible anatomical character differing from the latter genus, I do not see my way clear to make such a separation. Conchologically, such species as C. villanus Phil., are very near to T. scabrosus Phil., etc.; and finally, when we see that neither the teeth of the columella nor of the outer lip are constant in nearly all the species, nor the form of shell, peculiar umbilicus nor granulated sculpture can be depended on for generic characters, it is clear that the group can have no higher rank than Tectus, Infundibulum, and other subdivisions under Trochus.
As to the grouping of the species I have not been able to discover any mode of division into sections wholly satisfactory to myself. Perhaps the species are best grouped by a division into two sections ; Clanculus including those with strong basal tooth to the columella, and Clanculopsis for species with only a small denticle at the base. Each of these sections may be divided into species with the columella solute or tortuous above, inserted at or near the center of the axis in the bottom of the false umbilicus, and those with the columella inserted upon the edge of the umbilicus, and not deeply entering. Perhaps this last distinction is the more fundamental of the two. There are a number of species which greatly resemble species of Gibbula of the section Eurytrochus. Of course the resemblance is wholly superficial, as there is no real relationship between the forms.
Tryon, G.W. & Pilsbry, H.A.. Manual of Conchology; Vol. XI ; Trochidae
Subgenus 1 : Clanculus s.s.
The species included in this subgenus have the aperture with very well developed teeth; the basal tooth of the columella is complex, sculptured with two or more folds; on the inside of the outer lip, there are internal teeth extending into spiral lirae; the upper tooth is sometimes fused with the closer one, forming a large anal tooth.
Subgenus 2 : Clanculiopsis
The species of this subgenus have the apertural teeth weakly developed; the columella presents several simple small teeth, a little larger and more prominent than those at the base; anal tooth absent, because the top ridge inside the outer lip is neither enlarged nor fused with the next one.
Rubio, F. & Rolán, E., 2002. Revision of the Genus Clanculus (Gastropoda: Trochidae) in the Eastern Atlantic.
Author: Jan Delsing

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Included taxa

Number of records: 93

species Clanculus alarconi H.A. Rehder, 1980
species Clanculus albanyensis Jansen, 1995
species Clanculus albinus Adams, 1853

Clanculus albinus

species Clanculus albugo (Watson, 1880)
species Clanculus aloysii Tenison-Woods, 1876 - Saint Aloysius' Top-shell

Clanculus aloysii

species Clanculus amaeunus R. A. Philippi, 1852

Clanculus amaeunus

species Clanculus assabensis Caramagna, 1888

Clanculus assabensis

species Clanculus ater Pilsbry, 1901
species Clanculus atricatenus J. R. Tomlin, 1921

Clanculus atricatenus

species Clanculus atropurpureus (Gould, 1849)

Clanculus atropurpureus

species Clanculus atypicus Iredale, 1912

Clanculus atypicus

species Clanculus bathyraphe Smith, 1862

Clanculus bathyraphe

species Clanculus berthelotii (d'Orbigny, 1840)

Clanculus berthelotii

species Clanculus bicarinatus Angas, 1880

Clanculus bicarinatus

species Clanculus boyeti Poppe, Tagaro & Dekker, 2006

Clanculus boyeti

species Clanculus bronni R. W. Dunker, 1860

Clanculus bronni

species Clanculus brunneus Adams, 1853

Clanculus brunneus

species Clanculus buysei Poppe, Tagaro & Dekker, 2006

Clanculus buysei

species Clanculus ceylonicus Nevill & Nevill, 1869

Clanculus ceylonicus

species Clanculus clanguloides (Wood, 1828)

Clanculus clanguloides

species Clanculus clangulus (Wood, 1828)

Clanculus clangulus

species Clanculus cognatus (H.A. Pilsbry, 1903)

Clanculus cognatus

species Clanculus comarilis Hedley, 1912

Clanculus comarilis

species Clanculus consobrinus Tate, 1893 - Cingulate Top Shell

Clanculus consobrinus

species Clanculus corallinus (Gmelin, 1791)

Clanculus corallinus

species Clanculus cristinae Rubio & Rolán, 2002

Clanculus cristinae

species Clanculus cruciatus (Linnaeus, 1758)

Clanculus cruciatus

species Clanculus denticulatus (Gray, 1826) - Toothed Top Shell

Clanculus denticulatus

species Clanculus depictus A. Adams, 1855

Clanculus depictus

species Clanculus dunkeri (Koch in Philippi, 1843) - Dunker's Top-shell

Clanculus dunkeri

species Clanculus edentulus A. Adams, 1853
species Clanculus escondidus Poppe, Tagaro & Vilvens, 2009

Clanculus escondidus

species Clanculus euchelioides Tate, 1893

Clanculus euchelioides

species Clanculus festivus Tapparone Canefri, 1874 inq.

Clanculus festivus

species Clanculus flagellatus (Philippi, 1849) - Flamed Top-shell

Clanculus flagellatus

species Clanculus floridus (Philippi, 1850)

Clanculus floridus

species Clanculus flosculus P. Fischer, 1880

Clanculus flosculus

species Clanculus gemmulifer H. A. Pilsbry, 1901
species Clanculus granoliratus T. A. di Monterosato, 1889

Clanculus granoliratus

species Clanculus granti Hedley, 1907

Clanculus granti

species Clanculus guineensis (Gmelin, 1791)

Clanculus guineensis

species Clanculus hizenensis H.A. Pilsbry, 1901

Clanculus hizenensis

species Clanculus johnstoni Hedley, 1917

Clanculus johnstoni

species Clanculus jucundus Gould, 1861
species Clanculus jussieui (Payraudeau, 1826)

Clanculus jussieui

species Clanculus kinzelbachi F. Nordsieck, 1982
species Clanculus korkosi Singer, Mienis & Geiger, 2000

Clanculus korkosi

species Clanculus kraussi Philippi in Martini & Chemnitz, 1849

Clanculus kraussi

species Clanculus largillierti (Philippi, 1849)

Clanculus largillierti

species Clanculus laurae Cecalupo, Buzzurro & Mariani, 2008
species Clanculus leucomphalus Verco, 1905 - White Margined Top Shell

Clanculus leucomphalus

species Clanculus limbatus (Quoy & Gaimard, 1834) - Keeled Top-shell

Clanculus limbatus

species Clanculus margaritarius (Philippi, 1846) - Beautiful Clanculus

Clanculus margaritarius

species Clanculus mariaemaris Rubio & Rolán, 2002

Clanculus mariaemaris

species Clanculus maugeri (Wood, 1828)

Clanculus maugeri

species Clanculus mauritianus Melvill, 1909

Clanculus mauritianus

species Clanculus maxillatus (Menke, 1843) - Dotted Top Shell

Clanculus maxillatus

species Clanculus microdon A. Adams, 1853

Clanculus microdon

species Clanculus miniatus H. E. Anton, 1839

Clanculus miniatus

species Clanculus minor A. Adams, 1853

Clanculus minor

species Clanculus mixtus E. A. Smith, 1903

Clanculus mixtus

species Clanculus multipunctatus Jansen, 1995
species Clanculus natalensis Herbert, 1993

Clanculus natalensis

species Clanculus nexus Bozzetti, 2014
species Clanculus ochroleucus (Philippi, 1853) - Yellow Top Shell

Clanculus ochroleucus

species Clanculus peccatus Finlay, 1927
species Clanculus persicus T. Habe & Shikama, 1964

Clanculus persicus

species Clanculus personatus (Philippi, 1849) - Constricted Top Shell

Clanculus personatus

species Clanculus petziae Rubio, Rolán & Ryall, 2002

Clanculus petziae

species Clanculus pharaonius (Linnaeus, 1758) - Strawberry Top Shell

Clanculus pharaonius

species Clanculus philippi (Koch in Philippi, 1843) - Philippi's Top Shell

Clanculus philippi

species Clanculus pini Rubio & Rolán, 2002

Clanculus pini

species Clanculus plebejus (Philippi, 1851) - People's Top-shell

Clanculus plebejus

species Clanculus pseudocorallinus Gofas, 1984

Clanculus pseudocorallinus

species Clanculus puniceus (Philippi, 1846)

Clanculus puniceus

species Clanculus rarus (Dufo, 1840)
species Clanculus relictus S. Kosuge & Koizumi, 1990
species Clanculus richeri Vilvens, 2000
species Clanculus ringens (Menke, 1843) - Ridged Top Shell

Clanculus ringens

species Clanculus samoensis (Hombron & Jacquinot, 1848)
species Clanculus santamariae Gofas, 1984

Clanculus santamariae

species Clanculus scabrosus (Philippi, 1850)

Clanculus scabrosus

species Clanculus scotti Poppe, Tagaro & Dekker, 2006

Clanculus scotti

species Clanculus simoni Poppe, Tagaro & Dekker, 2006

Clanculus simoni

species Clanculus spadiceus Philippi, 1848

Clanculus spadiceus

species Clanculus stigmatarius A. Adams, 1853

Clanculus stigmatarius

species Clanculus thomasi (Crosse, 1862)

Clanculus thomasi

species Clanculus tonnerrei (Nevill & Nevill, 1874)

Clanculus tonnerrei

species Clanculus undatoides Tenison-Woods, 1879

Clanculus undatoides

species Clanculus undatus (Lamarck, 1816) - Wavy Top-shell

Clanculus undatus

species Clanculus villanus (Philippi, 1846)

Clanculus villanus

species Clanculus waltonae G. B. Sowerby III, 1892

Clanculus waltonae

species Clanculus weedingi Cotton, 1939

Clanculus weedingi


Links and literature

EN The Interim Register of Marine and Nonmarine Genera [1062644]

Rees, T. (compiler): The Interim Register of Marine and Nonmarine Genera [http://www.irmng.org] [as Euriclanculus Cotton & Godfrey, 1934]
Data retrieved on: 8 December 2019
EN The Interim Register of Marine and Nonmarine Genera [1126421]

Rees, T. (compiler): The Interim Register of Marine and Nonmarine Genera [http://www.irmng.org] [as Isoclanculus Cotton & Godfrey, 1934]
Data retrieved on: 8 December 2019
EN The Interim Register of Marine and Nonmarine Genera [1167797]

Rees, T. (compiler): The Interim Register of Marine and Nonmarine Genera [http://www.irmng.org] [as Microclanculus Cotton & Godfrey, 1934]
Data retrieved on: 8 December 2019
EN The Interim Register of Marine and Nonmarine Genera [1189444]

Rees, T. (compiler): The Interim Register of Marine and Nonmarine Genera [http://www.irmng.org] [as Euclanculus Cotton & Godfrey, 1934]
Data retrieved on: 8 December 2019
EN The Interim Register of Marine and Nonmarine Genera [1192729]

Rees, T. (compiler): The Interim Register of Marine and Nonmarine Genera [http://www.irmng.org] [as Mesoclanculus Iredale, 1924]
Data retrieved on: 8 December 2019
EN The Interim Register of Marine and Nonmarine Genera [1261802]

Rees, T. (compiler): The Interim Register of Marine and Nonmarine Genera [http://www.irmng.org] [as Paraclanculus Finlay, 1926]
Data retrieved on: 8 December 2019
EN The Interim Register of Marine and Nonmarine Genera [1295567]

Rees, T. (compiler): The Interim Register of Marine and Nonmarine Genera [http://www.irmng.org] [as Clanculus Montfort, 1810]
Data retrieved on: 8 December 2019
EN Australian Faunal Directory [806d8969-a7c3-4318-a79c-d4cc95b6db79]

ABRS (2009-2019): Australian Faunal Directory [https://biodiversity.org.au/afd/home], Australian Biological Resources Study, Canberra [as Clanculus Montfort, 1810]
Data retrieved on: 15 February 2015

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Comments - 1


Explanations

inq. nomen inquirendum - doubtful identity, needs further investigation