SW Costa Rica, Panama (including Isla del Rey, Isla Contadora, Isla de Cébaco, Isla Suscantupu), Colombia (east of the Andes and north of the cordilleras Central and Oriental), N Venezuela (north of the Cordillera de Mérida and in the drainage of the Río Orinoco, north and west of the Guiana Shield, east to the Orinoco delta, Trujilllo